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First, tumor cells often exhibit mitochondrial dysfunction due to the limited repair capacity of mitochondrial DNA, which lacks introns and protective histones, coupled with a high mutation rate, leading to impaired mitochondrial function ( 2.2 Lipid metabolism Tumor cells also upregulate de novo fatty acid synthesis to support membrane biosynthesis and energy storage by increasing the expression of key enzymes such as fatty acid synthase (FASN) and acetyl-CoA carboxylase (ACC) ( 2.3 Amino acid metabolism Amino acid metabolism, especially glutamine metabolism, is pivotal for tumor cell proliferation, migration, and invasion

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